Garfield is a fat and free-spirited orange cat who lives with his owner, Jon Arbuckle. Garfield passes his time by antagonizing Jon and teasing an aggressive neighbor Doberman Pinscher, Luca. Aside from Jon, Garfield maintains an unlikely friendship with a helpful mouse, Louis. He also socializes with his fellow neighborhood cats, including Garfield's stooge Nermal and Arlene.
Meanwhile, a local television host, Happy Chapman, known for his cat "Persnikitty" is introduced as supposedly a happy man. In reality he is allergic to cats, jealous of his brother Walter J. Chapman, a news reporter, and to be more successful by performing on TV show Good Day New York. Jon has made a habit of bringing Garfield to the veterinarian, in-order to see vet Dr. Liz Wilson. Jon tries to ask her out, but due to a misunderstanding, he is given custody of a stray dog, Odie. Regardless, Jon and Liz begin dating. Garfield is angry at having to share the house with a dog, of whom Jon grows fond. Odie is brought to a canine talent show, where Liz is a judge. Garfield gets involved in an altercation there with other animals, which moves Odie to the center of the ring, where he begins dancing to "Hey Mama" by the Black Eyed Peas.
His improvised performance is a hit. Happy Chapman, who also is a judge of the dog show is impressed with Odie, and offers Jon a television deal for Odie, but Jon declines. After Garfield causes a mess inside Jon's house out of jealousy, Jon punishes Garfield by making him sleep outside for one night. Odie comes out to comfort Garfield but Garfield locks Odie out. Nermal and Arlene witness this as Odie runs away; he is then picked up by an elderly woman named Mrs. Baker. Jon searches with Liz for Odie while the neighborhood animals treat Garfield as an outcast. Meanwhile, Chapman and his assistant Wendell find a notice Mrs. Baker created of Odie and, recognizing the lucrative possibilities, claim Odie as Happy's own.
When Garfield sees Odie on television and hears Chapman announce he and Odie are going to New York City, Garfield sets out to rescue Odie. Jon discovers Garfield missing so Jon and Liz start searching. Garfield gets into the broadcast tower via the air vents but he is blown around violently. Garfield finds Odie locked in a room; Chapman enters and secures a shock collar to Odie, which, when activated, releases an electric discharge that forces him to perform tricks.
Chapman heads for the train station with Garfield in close pursuit. However, an animal control officer catches Garfield mistaking him as a runaway. Mrs. Baker tells Jon that Chapman took Odie, but Jon believes Garfield was taken too and he and Liz race to Telegraph Tower and then to the train station, after learning Chapman has left. Garfield is released from the pound by Chapman's abandoned feline star, Persnikitty, who is really named Sir Roland. Chapman boards a Texas-bound train, with Odie in the luggage car. Garfield arrives only to see the train depart. Garfield sneaks into the train system control room and rearranges some tracks, leading to an impending train wreck. Garfield hits an emergency control and causes Chapman/Odie's train to return to the station. Garfield frees Odie and they exit the train. However, Chapman chases them. Chapman threatens Odie with the shock collar, but is stopped by Garfield's friends and animals from the pound, led by Sir Roland. They swarm and attack Chapman, allowing Odie and Garfield to escape.
The shock collar is now on Chapman who gets shocked. Jon and Liz arrive to reclaim the animals and find Chapman disoriented. Jon punches Chapman in the face for stealing his pets, and leaves with Liz and the two animals. Chapman is arrested for his supposed involvement with the trains, as well as for abducting Odie. Garfield regains the respect of his animal friends as a hero. Back at home, Liz and Jon form a relationship, while Garfield learns a lesson about friendship.
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