In Alberta, Canada's Jasper National Park, Kate begins Alpha school
with her father and grows up as a fully trained Alpha, but on her first hunt
she crosses paths with two wolves from the Eastern pack who are lacking food in
their own territory and nearly start a war. The two packs have had a bitter
rivalry ever since Tony commanded the Eastern wolves to cross over into Western
territory (against pack law) to get sufficient food. Winston and Tony meet up
one night and arrange for Kate to marry Garth (Chris
Carmack), Tony's son. Kate, having a sense of responsibility and duty,
agrees for the good of the pack and meets Garth at the Moonlight Howl.
Garth seems perfect except that he has a terrible howl.
Kate, taken aback, leaves and has a talk with Humphrey about Garth, when
suddenly they are tranquilized by some park rangers and taken away to Sawtooth National Recreation Area in Idaho, where they
meet golfer Marcel and his caddy Paddy. Humphrey and Kate learn that they were
relocated to repopulate the species. This horrifies Kate, but intrigues
Humphrey so that he can be with Kate, but after coming to an agreement, Marcel
and Paddy help the wolves return home to Jasper Park.
The two packs discover that Kate has gone missing. Tony
warns Winston that on the next full moon if Kate doesn't return to marry Garth,
war will be declared for their territory. Kate's sister Lilly however has
developed a crush on Garth and suggests that she show him around.
Kate and Humphrey's ride home is cut short at a gas station,
where Humphrey is mistaken for a rabid wolf after he eats a cream-filled
cupcake. Kate attacks a man with the gun, and the two make flee into the
forest. During their homeward journey, they cross a dangerous ravine with
Humphrey saving Kate from a nasty fall.
The next morning, Marcel and Paddy find the wolves and
direct them to a train over a snowy mountain that will take them to Jasper.
Climbing the mountain, Humphrey meets agrizzly
bear cub whose never seen wolves, but a mishap gets the cub hurt
angering a trio of adult bears. Kate comes to Humphrey's rescue and after a
rough slide down the mountain manage to board the passing train.
During this time, Lilly and Garth are bonding. At the
midnight howl, Lilly compassionately teaches Garth how to howl effectively. At
the same time, Humphrey and Kate begin to fall in love. By this time, the full
moon has come and the both packs declare war against each other. As the train
passes by Jasper, Kate and Humphrey exchange of feelings is cut short by the
sight of the wolf pack war. Kate stops the fight by declaring she will marry
Garth. The next day, a heartbroken Humphrey bids his goodbyes with Kate,
Marcel, and Paddy deciding to leave by himself and Kate also feels the same
way.
During the ceremony, Kate decides not to marry Garth,
declaring her love for Humphrey and Garth declares his own romantic
love for Lilly. An Alpha wolf marrying an Omega wolf goes against pack
customs and a conflict ensues between the packs, but is cut short when a
massive stampede of caribou comes. Winston and Tony get caught in the stampede
but Kate and Humphrey rush in to save them. Kate is almost trampled but
survives thanks to Humphrey's protection. Later at the Moonlight Howl, all of
the wolves celebrate the love of Kate to Humphrey, and Garth to Lilly, breaking
the social classes and traditions. Humphrey and Kate sing a duet.
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